Rules for Authors


Arctoa is publishing papers on mosses, liverworts and hornworts. Printed and electronic versions at are issued.

The submitted information should be significant and novel, not published earlier and not submitted or considered for publications elsewhere.

Papers must be in English, except the sections of New Records and Regional Floras, where both English and Russian are acceptable. Further instruction for New Bryophyte Records and Regional Floras sections is given below.




Manuscripts should be submitted into two addresses:

Optimally to name files with the surname of the first author (e.g., ivanov-text.rtf, ivanov-fig1.tif, etc.) written by letters of the Latin alphabet.

Manuscripts must be fully prepared for publication, checked for language and the following rules. Numerous technical and orthographic errors preclude sending paper to reviewers, authors will be requested to improve and resubmit their papers. English editing is a responsibility of author(s).


Titles, author names and abstracts will be published in both English and Russian. Authors can provide their own Russian version; however, if this information will not be supplied, then it will be completed by the editor.

1. Text of the manuscript includes: Title, Author(s), Author address and e-mail, Abstract, Text of paper, Literature Cited, Figure explanations, Table explanations, Tables. Files in *.doc and *.rtf are acceptable. In titles and authors do not use heading letters (except the first letter, as in usual text). Please suggest running head if the paper title is long; in the absence of suggestion, the editor can use his own version of the running head.

File with manuscript text must not include Figures. Each figure must be submitted as a separate file with indication in file name fig-1, fig-2, etc. Files in *.tif, *.jpg, and *.eps are acceptable and authors are requested to convert other files to these formats. DO NOT send figures embedded in files of text editors, e.g. MSWord. In case of difficulties, please, consult the editor for the possible exceptions. Line drawings should be preferably in black and white bitmap mode (no less 600 dpi, 1200 dpi recommended) or in grayscale mode (no less 200 dpi, 300-600 dpi recommended). Avoid figures in indexed colors. Figures must be of good quality and should have scale bars. In case of photographs, author can suggest the bar of certain length (preferably) rather than insert the bar himself. They can be arranged in plates or given separately.

4. Color photographs may be printed at author expense. However, color image files (RGB mode) that can be printed clearly in grayscale mode can be submitted as well, with indication that they are to be printed in grayscale mode; those figures will be in color in pdf file. In these cases authors should submit two files: color of grayscale ones, with the same beginning of name, e.g. ivanov-fig1-rgb.tif (or ivanov-fig1-color.tif) AND ivanov-fig1-black-white.tif.

Previously published figures will be accepted only with the writing permission of the copyright holder; author are requested to obtain these permissions and submit to Arctoa along with the manuscript.


Footnotes should be avoided as much as possible; each footnote has to be submitted as a separate file (and text should have reference to this file in appropriate place, FOOTNOTE1, etc.). This does not concern author addresses that are given immediately after authors of the paper.

After paper acceptance for publications, author must sign copyright agreement, transferring copyright to Arctoa. At the same time, authors will have all the rights for the subsequent use of their data, referring to the original publication. Copyright agreement can be taken from




1. Only SI units must be used.

2. The Latin names of species and infraspecific taxa should include author(s) at first mention. However, if the paper has a list or catalogue of all species, then authors should be mentioned in that list and not in the text. If species concepts follow one of checklists and all the species names would have the species authors as in that checklist, then please give reference to this checklist and do not provide any authors of species and infraspecific taxa. Author abbreviations should follow

3. Herbarium acronyms must follow Index Herbariorum,




1. The Latin names of plants and animals must be in italics for taxa of rank of genera and below (e.g. genera, species, infraspecific taxa), but in normal font for taxa above genera (e.g. families, classes, etc.); rank of taxa (e.g. var., subsp.) in normal font), No other formatting (font, paragraph, style) in text is needed, except letters absent in English alphabet (see below) and literature references (as explained below).

2. Microns should be given as mkm. Letters ä ü ö should be given as ae, ue, oe respectively and underlined. Other specific letters should be underlined, but their correct printing is not guaranteed.

3. Dates should be given as follow: day (in Arabic numerals).month (in Roman numerals).year (in Arabic, in four numerals), e.g. 25.VII.1998

4. Decimal fraction should be written with point (0.1), not comma (0,1).

5. Variation in dimensional characters should be given with hyphen, (20-)25-30, not with dash; note that no space should be near hyphen in this case.

6. Dash can be given as hyphen or dash, but always with space before and after it.

7. Space must be always put after numerals, point (except in decimal and dates), comma, colon, semicolon. Never put two spaces together.

8. Reference to figures should be numbered. If the paper has only one figure it must be Fig. 1. All figures has to be discussed in the text.

9. References in text must be as follow: Ivanov (2000), or (Ivanov, 2000); Ivanov & Sidorov (2000), or (Ivanov & Sidorov, 2000); Ivanov et al. (2000), or (Ivanov et al., 2000).

10. Literature citation must include authors names written in ordinary way, with the only first heading letter; All-caps style will be given in the course of the final technical preparation:

POTEMKIN, A.D. 2001. Three new species of Scapania (Hepaticae) from India and China. Ann. Bot. Fennici 38: 1-7.

[BARDUNOV, L.V.] , .. 1999. . [On the rare species in the moss flora of Central Siberia] . . [Bot. Zhurn.] 84(2): 73-78.

[BELKINA, O.A. & A.Yu. LIKHACHEV] , .., .. 1999. . [To cliff moss flora of Murmansk Region mountains (North-West Russia)] : (. , ..) , [In: Konstantinova, N.A. Flora i rastitelnost Murmanskoj oblasti. Apatity, Kolskij nauchnyj zentr Ross. Akad. Nauk]: 31-54.

Smith, A. J. E. 1978. The moss flora of Britain and Ireland. Cambridge & al., Cambridge Univ. Press. 706 pp.

[SAVICZ-LYUBITSKAYA, L.I. & Z.N. SMIRNOVA] -, .., .. 1970. . . [The handbook of mosses of the USSR. The acrocarpous mosses] ., [Leningrad, Nauka], 826 pp.

[BARDUNOV, L.V. & A.N. VASILYEV] , .., .. 1998. . [The bryoflora of forests of Siberia] .: XX-XXI . . ., II(X) (26-29 1998, -). . 2. : [In: Problemy botaniki na rubezhe XX-XXI vekov. Abstr. II(X) Congr. Russ. Bot. Soc. (26-29 May, 1998, St.Petersburg). Vol. 2. St.-Petersburg, Komarov' Bot. Inst.]: 127-128.


Latin names are given in normal font, not italics, in Literature Cited.

Journal, book, pages must be in italics. Volume must be boldfaced.

Order of literature references is alphabetical, following Latin transliteration of Cyrillic alphabet.




Manuscripts will be reviewed and authors will be informed of the overall evaluation of the submitted paper, and may be requested to complete, delete or amend part of paper. In case of disagreement with reviewer opinion, the author should provide explanation of his position to the editor. In case of major revision, the manuscript may be send to the additional review.


Proof reading copy


After paper acceptance, the proof reading copy will be send to author with request to check the paper and return corrected copy as soon as possible. The non-receipt of proof reading copy within ten days does not guaranteed that corrections will be included in publication. If paper has more than one author, then proof reading copy will be send to the first author unless otherwise is indicated. Numerous mistakes of original manuscript will be corrected in proof reading file only at author expense.




Printed version in *.pdf file will be provided to authors free of charge. No separate reprints will be provided.



New Bryophyte Records section

New Bryophyte Records section presents short notes about interesting records, as well as information about species missing for certain provinces and countries. Each record must have longitude and latitude data (if this is not indicated in label, please give this approximately [with ~] according to map). Name of collector should be in both Latin and Cyrillic alphabet. Collection date (prior the collector) and collector number (after collector name) should be also given, as well as herbarium where the specimen is deposited.



Since 21 volume, the format of New Bryophyte Records in Arctoa will be slightly changes. It will appear as one article of many authors, and (1) will have one bibliography for all contributions; (2) list of authors wih address will be after bibliography (as well as in earlier volumes); (3) ackowledgements will be restricted by abbreviation of foundation and and grant number, written after author address.

Citation of New Bryophyte Record material may be in two ways:

Ivanov I.I. 2020. New moss records from Ivanono Province. 1. In: Sofronova E.V. (ed.) New bryophyte records. 10. Arctoa 30: 300-350.

Sofronova E.V., Aleksandrov A.A., Ivanov A.I., Ivanov I.A., Ivanov I.I., Volkov V.V. 2020. New bryophyte records. 10. Arctoa 30: 300-350.

The series of New Bryophyte Records will start in vol. 21 since the first number: . 1 -- New bryophyte records. 1, however, the previous numbering from vols. 15-20 for separate regions will remain.



New moss records from Ivanono Province. 5. . 5. I.I. Ivanov ..

New for Russia

Ivanovia ivanovii Ivanov Ivanovo Distr., Ivanovka, (9393'N 18993'E), 14000 m alt., on limestone with Cephalozia bicuspidata, 32.V.2020 Ivanov # 666666 [] {IVAN}.



Regional Floras section

Section will publish papers with complete lists of moss/ hepatic floras from poorly known regions that have sufficient novelty for this region. Papers in Russian and English are accepted, in the former case a reader have to be able understand the study area and species frequency. Captions for tables and illustrations and abstract must be in two languages. Welcomed are the overviews of habitat types, including characteristic bryophytes.


New Publication


New Publication sections is publishing bibliography on bryological literature concerning East Europe and North Asia, and represented in at least one of the following libraries: of Komarov Botanical Institute in St.-Petersburg, Main Botanical Garden in Moscow, and Institute of Biology in Ufa, or pdf-file (if not from Journal, then with the title page), sent to three addresses: Irina Czernyadjeva <>, Elena Sofronova <>,